FORM 4
[ ] Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).         
 UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549

STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP OF SECURITIES
                                                                                  
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Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934 or Section 30(h) of the Investment Company Act of 1940
                      

1. Name and Address of Reporting Person *

WELLS FARGO & COMPANY/MN		 2. Issuer Name and Ticker or Trading Symbol

BLACKROCK MUNIHOLDINGS CALIFORNIA QUALITY FUND, INC. [ MUC ] 		5. Relationship of Reporting Person(s) to Issuer (Check all applicable)

_____ Director                    __X__ 10% Owner
_____ Officer (give title below)    _____ Other (specify below)
(Last)          (First)          (Middle)

420 MONTGOMERY STREET		 3. Date of Earliest Transaction (MM/DD/YYYY)

4/11/2022
(Street)

SAN FRANCISCO, CA 94104
(City)        (State)        (Zip)
4. If Amendment, Date Original Filed (MM/DD/YYYY)

 
 6. Individual or Joint/Group Filing (Check Applicable Line)

___ Form filed by One Reporting Person
_ X _ Form filed by More than One Reporting Person

Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned
1.Title of Security
(Instr. 3) 		2. Trans. Date 2A. Deemed Execution Date, if any 3. Trans. Code
(Instr. 8) 		4. Securities Acquired (A) or Disposed of (D)
(Instr. 3, 4 and 5) 		5. Amount of Securities Beneficially Owned Following Reported Transaction(s)
(Instr. 3 and 4) 		6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4) 7. Nature of Indirect Beneficial Ownership (Instr. 4)
Code V Amount (A) or (D) Price
Variable Rate Muni Term Preferred Shares 4/11/2022  J(1)  2724 A(1) (1)5264 (1)I (2)By Subsidiary (2)(3)

Table II - Derivative Securities Beneficially Owned (e.g., puts, calls, warrants, options, convertible securities)
1. Title of Derivate Security
(Instr. 3)		2. Conversion or Exercise Price of Derivative Security3. Trans. Date3A. Deemed Execution Date, if any4. Trans. Code
(Instr. 8)		5. Number of Derivative Securities Acquired (A) or Disposed of (D)
(Instr. 3, 4 and 5)		6. Date Exercisable and Expiration Date7. Title and Amount of Securities Underlying Derivative Security
(Instr. 3 and 4)		8. Price of Derivative Security
(Instr. 5)		9. Number of derivative Securities Beneficially Owned Following Reported Transaction(s) (Instr. 4)10. Ownership Form of Derivative Security: Direct (D) or Indirect (I) (Instr. 4)11. Nature of Indirect Beneficial Ownership (Instr. 4)
CodeV(A)(D)Date ExercisableExpiration DateTitleAmount or Number of Shares
Explanation of Responses:
(1) In connection with the reorganization of BlackRock MuniYield California Fund, Inc. ("MYC") and BlackRock MuniYield California Quality Fund, Inc. ("MCA") into the Issuer, 1,059 Series W-7 Variable Rate Muni Term Preferred Shares ("VMTP Shares") of MYC and 1,665 VMTP Shares of MCA beneficially owned by Wells Fargo Bank, N.A. ("WFBNA") were exchanged for an equal number of VMTP Shares of the Issuer in a cashless transaction. In addition to the VMTP Shares of MYC and MCA that were exchanged for an equal number of VMTP Shares of the Issuer, 2,540 VMTP Shares of the Issuer were previously owned by WFBNA and reported on that cetain Form 3 dated and filed as of March 29, 2012, which form is updated by the filing of this Form 4. WFBNA is a wholly owned subsidiary of Wells Fargo & Company ("Wells Fargo").
(2) This statement is jointly filed by Wells Fargo and WFBNA. Wells Fargo holds an indirect interest in the securities listed in Table I by virtue of its indirect ownership of its subsidiary WFBNA.
(3) Each reporting person declares that neither the filing of this statement nor anything herein shall be construed as an admission that such person is, for the purposes of Section 13(d) of the US Securities Exchange Act of 1934 or any other purpose, (i) acting (or has agreed or is agreeing to act together with any other person) as a partnership, limited partnership, syndicate or other group for the purpose of acquiring, holding or disposing of securities of the Issuer or otherwise with respect to the Issuer or any securities of the Issuer or (ii) a member of any group with respect to the Issuer or any securities of the Issuer.

Remarks:
Exhibits Index
Exhibit 99.1 - Joint Filing Agreement
Exhibit 99.2 - Joint Filer Information

Reporting Owners
Reporting Owner Name / Address
Relationships
Director10% OwnerOfficerOther
WELLS FARGO & COMPANY/MN
420 MONTGOMERY STREET
SAN FRANCISCO, CA 94104
X

WELLS FARGO BANK N A
101 NORTH PHILLIPS AVENUE
SIOUX FALLS, SD 57104
X
Signatures
WELLS FARGO & COMPANY, by: /s/ Patricia Arce4/13/2022
**Signature of Reporting PersonDate

WELLS FARGO BANK, N.A., by: /s/ Alejandro Piekarewicz4/13/2022
**Signature of Reporting PersonDate
Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
*If the form is filed by more than one reporting person, see Instruction 4(b)(v).
**Intentional misstatements or omissions of facts constitute Federal Criminal Violations. See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
Note:File three copies of this Form, one of which must be manually signed. If space is insufficient, see Instruction 6 for procedure.
Persons who respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB control number.
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